Advances in Legume Systematics 14. Classification of Caesalpinioideae. Part 2: Higher-level classification
Por:
Bruneau A., De Queiroz L.P., Ringelberg J.J., Borges L.M., Bortoluzzi R.L.D.C., Brown G.K., Cardoso D.B.O.S., Clark R.P., Conceição A.D.S., Cota M.M.T., Demeulenaere E., De Stefano R.D., Ebinger J.E., Ferm J., Fonseca-Cortés A., Gagnon E., Grether R., Guerra E., Haston E., Herendeen P.S., Hernández H.M., Hopkins H.C.F., Huamantupa-Chuquimaco I., Hughes C.E., Ickert-Bond S.M., Iganci J., Koenen E.J.M., Lewis G.P., De Lima H.C., De Lima A.G., Luckow M., Marazzi B., Maslin B.R., Morales M., Morim M.P., Murphy D.J., O’Donnell S.A., Oliveira F.G., Oliveira A.C.D.S., Rando J.G., Ribeiro P.G., Ribeiro C.L., Santos F.D.S., Seigler D.S., da Silva G.S., Simon M.F., Soares M.V.B., Terra V.
Publicada:
1 ene 2024
Resumen:
Caesalpinioideae is the second largest subfamily of legumes (Leguminosae) with ca. 4680 species and 163 genera. It is an ecologically and economically important group formed of mostly woody perennials that range from large canopy emergent trees to functionally herbaceous geoxyles, lianas and shrubs, and which has a global distribution, occurring on every continent except Antarctica. Following the recent re-circumscription of 15 Caesalpinioideae genera as presented in Advances in Legume Systematics 14, Part 1, and using as a basis a phylogenomic analysis of 997 nuclear gene sequences for 420 species and all but five of the genera currently recognised in the subfamily, we present a new higher-level classification for the subfamily. The new classification of Caesalpinioideae comprises eleven tribes, all of which are either new, reinstated or re-circumscribed at this rank: Caesalpinieae Rchb. (27 genera / ca. 223 species), Campsiandreae LPWG (2 / 5-22), Cassieae Bronn (7 / 695), Cera-tonieae Rchb. (4 / 6), Dimorphandreae Benth. (4 / 35), Erythrophleeae LPWG (2 /13), Gleditsieae Nakai (3 / 20), Mimoseae Bronn (100 / ca. 3510), Pterogyneae LPWG (1 / 1), Schizolobieae Nakai (8 / 42-43), Sclerolobieae Benth. & Hook. f. (5 / ca. 113). Although many of these lineages have been recognised and named in the past, either as tribes or informal generic groups, their circumscriptions have varied widely and changed over the past decades, such that all the tribes described here differ in generic membership from those previously recognised. Importantly, the approximately 3500 species and 100 genera of the former subfamily Mimosoideae are now placed in the reinstated, but newly circumscribed, tribe Mimoseae. Because of the large size and ecological importance of the tribe, we also provide a clade-based classification system for Mimoseae that includes 17 named lower-level clades. Fourteen of the 100 Mimoseae genera remain unplaced in these lower-level clades: eight are resolved in two grades and six are phylogenetically isolated monogeneric lineages. In addition to the new classification, we provide a key to genera, morphological descriptions and notes for all 163 genera, all tribes, and all named clades. The diversity of growth forms, foliage, flowers and fruits are illustrated for all genera, and for each genus we also provide a distribution map, based on quality-controlled herbarium specimen localities. A glossary for specialised terms used in legume morphology is provided. This new phylogenetically based classification of Caesalpinioideae provides a solid system for communication and a framework for downstream analyses of biogeography, trait evolution and diversification, as well as for taxonomic revision of still understudied genera. © 2024 Anne Bruneau et al. This is an open access article distributed under terms of the Creative Commons Attribution License (Attribution 4.0 International -CC BY 4.0). All Rights Reserved.
Filiaciones:
Bruneau A.:
Institut de recherche en biologie végétale and Département de Sciences biologiques, Université de Montréal, 4101 Sherbrooke E., Montreal, H1X 2B2, QC, Canada
De Queiroz L.P.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Ringelberg J.J.:
Department of Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, Zurich, 8008, Switzerland
School of Geosciences, University of Edinburgh, Old College, South Bridge, Edinburgh, EH8 9YL, United Kingdom
Borges L.M.:
Universidade Federal de São Carlos, Departamento de Botânica, Rodovia Washington Luís, Km 235, 13565-905, SP, São Carlos, Brazil
Bortoluzzi R.L.D.C.:
Programa de Pós-graduação em Produção Vegetal, Universidade do Estado de Santa Catarina, Centro de Ciências Agroveterinárias, Avenida Luiz de Camões 2090, Lages, Santa Catarina, 88520-000, Brazil
Brown G.K.:
Queensland Herbarium and Biodiversity Science, Department of Environment and Science, Toowong, 4066, QLD, Australia
Cardoso D.B.O.S.:
Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Pacheco Leão 915, RJ, Rio de Janeiro, 22460-030, Brazil
Programa de Pós-Graduação em Biodiversidade e Evolução (PPGBioEvo), Instituto de Biologia, Universidade Federal de Bahia (UFBA), Rua Barão de Jeremoabo, s.n., Ondina, Salvador, BA, 40170-115, Brazil
Accelerated Taxonomy Department, Royal Botanic Gardens, Kew, Richmond, TW9 3AE, United Kingdom
Programa de Pós-graduação em Diversidade Vegetal, Universidade do Estado da Bahia, Herbário HUNEB, Campus VIII, Rua do Gangorra 503, Paulo Afonso, Bahia, 48608-240, Brazil
Center for Island Sustainability and Sea Grant, University of Guam, UOG Station, Mangilao, 96923, Guam
Centro de Investigación Científica de Yucatán, A.C. (CICY), Calle 43 No. 130 x 32 y 34, Chuburná de Hidalgo, CP 97205, Yucatán, Mérida, Mexico
Department of Ecology, Environment and Plant Sciences, Stockholm University, Stockholm, 10691, Sweden
Royal Botanic Garden Edinburgh, 20A Inverleith Row, Edinburgh, EH3 5LR, United Kingdom
Departamento de Biología, Universidad Autónoma Metropolitana-Iztapalapa, Apdo. Postal 55-535, Ciudad de México, 09340, Mexico
Chicago Botanic Garden, 1000 Lake Cook Road, Glencoe, 60022, IL, United States
Departamento de Botánica, Instituto de Biología, Universidad Nacional Autónoma de México, Cd. Universitaria, Ciudad de México, 04510, Mexico
Herbario Alwyn Gentry (HAG), Universidad Nacional Amazónica de Madre de Dios (UNAMAD), AV. Jorge Chávez N°1160, Madre de Dios, Peru
Department of Biology & Wildlife & Herbarium (ALA), University of Alaska Museum of the North, University of Alaska Fairbanks, P.O. Box 756960, Fairbanks, 99775-6960, AK, United States
Programa de Pós-Graduação em Fisiologia Vegetal, Universidade Federal de Pelotas, Instituto de Biologia, Campus Universitário Capão do Leão, Passeio André Dreyfus, Departamento de Botânica, Prédio 21, Rio Grande do Sul, Pelotas, 96010-900, Brazil
Evolutionary Biology & Ecology, Université Libre de Bruxelles, Faculté des Sciences, Campus du Solbosch - CP 160/12, Avenue F.D. Roosevelt, 50, Bruxelles, 1050, Belgium
Instituto Nacional da Mata Atlântica, INMA-MCTI, Av. José Ruschi, 4, Centro, 29650-000, Santa Teresa, Espírito Santo, Brazil
Department of Biological and Environmental Sciences, University of Gothenburg, Gothenburg, Sweden
School of Integrative Plant Science, Plant Biology Section, Cornell University, 215 Garden Avenue, Roberts Hall 260, Ithaca, 14853, NY, United States
Natural History Museum of Canton Ticino, Viale C. Cattaneo 4, Lugano, 6900, Switzerland
Singapore Herbarium, 1 Cluny Road, Singapore, Singapore
Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Godoy Cruz 2290 (C1425FQB), Ciudad Autónoma de Buenos Aires, Argentina
Royal Botanic Gardens Victoria, Melbourne, 3004, VIC, Australia
Geography and Environmental Sciences, Northumbria University, Ellison Place, Newcastle upon Tyne, NE1 8ST, United Kingdom
Programa de Pós-graduação em Ciências Ambientais, Universidade Federal do Oeste da Bahia, Rua Professor José Seabra Lemos 316, Bahia, Barreiras, 47800-021, Brazil
Department of Plant Biology, University of Illinois, Urbana, 61801, IL, United States
Instituto de Biologia, Universidade Estadual de Campinas, Campinas, SP, São Paulo, 13083-876, Brazil
Empresa Brasileira de Pesquisa Agropecuária (Embrapa), Recursos Genéticos e Biotecnologia, Parque Estação Biológica, Caixa Postal 02372, DF, Brasília, 70770-917, Brazil
Instituto de Biologia, Universidade Federal de Santa Maria, Santa Maria/RS, 97105-900, Brazil
Clark R.P.:
Institut de recherche en biologie végétale and Département de Sciences biologiques, Université de Montréal, 4101 Sherbrooke E., Montreal, H1X 2B2, QC, Canada
Conceição A.D.S.:
Institut de recherche en biologie végétale and Département de Sciences biologiques, Université de Montréal, 4101 Sherbrooke E., Montreal, H1X 2B2, QC, Canada
Cota M.M.T.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Demeulenaere E.:
Institut de recherche en biologie végétale and Département de Sciences biologiques, Université de Montréal, 4101 Sherbrooke E., Montreal, H1X 2B2, QC, Canada
De Stefano R.D.:
Institut de recherche en biologie végétale and Département de Sciences biologiques, Université de Montréal, 4101 Sherbrooke E., Montreal, H1X 2B2, QC, Canada
Ebinger J.E.:
Eastern Illinois University, Charleston, 61920, IL, United States
Ferm J.:
Institut de recherche en biologie végétale and Département de Sciences biologiques, Université de Montréal, 4101 Sherbrooke E., Montreal, H1X 2B2, QC, Canada
Fonseca-Cortés A.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Gagnon E.:
Institut de recherche en biologie végétale and Département de Sciences biologiques, Université de Montréal, 4101 Sherbrooke E., Montreal, H1X 2B2, QC, Canada
Department of Integrative Biology, University of Guelph, 50 Stone Road, Guelph, N1G 2W1, ON, Canada
Chair of Phytopathology, Technical University Munich, Freising, 85354, Germany
Grether R.:
Institut de recherche en biologie végétale and Département de Sciences biologiques, Université de Montréal, 4101 Sherbrooke E., Montreal, H1X 2B2, QC, Canada
Guerra E.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Haston E.:
Institut de recherche en biologie végétale and Département de Sciences biologiques, Université de Montréal, 4101 Sherbrooke E., Montreal, H1X 2B2, QC, Canada
Herendeen P.S.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Hernández H.M.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Hopkins H.C.F.:
Institut de recherche en biologie végétale and Département de Sciences biologiques, Université de Montréal, 4101 Sherbrooke E., Montreal, H1X 2B2, QC, Canada
Huamantupa-Chuquimaco I.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Hughes C.E.:
Department of Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, Zurich, 8008, Switzerland
Ickert-Bond S.M.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Iganci J.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Universidade Federal do Rio Grande do Sul, Programa de Pós-Graduação em Botânica, Av. Bento Gonçalves 9500, Bloco IV - Prédio 43433, RS, Porto Alegre, 91501-970, Brazil
Koenen E.J.M.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Lewis G.P.:
Institut de recherche en biologie végétale and Département de Sciences biologiques, Université de Montréal, 4101 Sherbrooke E., Montreal, H1X 2B2, QC, Canada
De Lima H.C.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Pacheco Leão 915, RJ, Rio de Janeiro, 22460-030, Brazil
De Lima A.G.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Pacheco Leão 915, RJ, Rio de Janeiro, 22460-030, Brazil
Luckow M.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Marazzi B.:
Department of Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, Zurich, 8008, Switzerland
Maslin B.R.:
Department of Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, Zurich, 8008, Switzerland
Western Australian Herbarium, Department of Biodiversity, Conservation and Attractions, Locked Bag 104, Bentley Delivery Centre, 6983, WA, Australia
Morales M.:
Department of Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, Zurich, 8008, Switzerland
Instituto de Recursos Biológicos, CIRN-CNIA, INTA. N. Repetto & Los Reseros s.n., Hurlingham, Buenos Aires, Argentina
Morim M.P.:
Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Pacheco Leão 915, RJ, Rio de Janeiro, 22460-030, Brazil
Murphy D.J.:
Department of Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, Zurich, 8008, Switzerland
O’Donnell S.A.:
Department of Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, Zurich, 8008, Switzerland
Oliveira F.G.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Oliveira A.C.D.S.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Rando J.G.:
Department of Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, Zurich, 8008, Switzerland
Ribeiro P.G.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Ribeiro C.L.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Santos F.D.S.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Seigler D.S.:
Department of Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, Zurich, 8008, Switzerland
da Silva G.S.:
Department of Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, Zurich, 8008, Switzerland
Simon M.F.:
School of Geosciences, University of Edinburgh, Old College, South Bridge, Edinburgh, EH8 9YL, United Kingdom
Soares M.V.B.:
Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Av. Transnordestina s/n, Campus, Novo Horizonte. 44036-900, BA, Feira de Santana, Brazil
Terra V.:
School of Geosciences, University of Edinburgh, Old College, South Bridge, Edinburgh, EH8 9YL, United Kingdom
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